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Captain Jack Harkness, played by Scottish-born American actor John Barrowman, is a member of Torchwood Three. A former Time Agent born in the 51st century, Jack had two years of his memories wiped by the Time Agency and subsequently became a con-man operating in the 20th century, where he stole the name of "Captain Jack Harkness"; his real name is unknown. After contact with the Ninth and Tenth Doctors in Doctor Who, Jack reformed and changed his ways to do good works with Torchwood where he has become a member of the Institute's Cardiff branch. He joined Torchwood in the 19th Century by time-travelling there. As a result of his reluctant immortality, Jack has lived a long life, in which he has seen the loss of his brother Gray, countless deaths and many different lovers over the years, both men and women.

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Gwen Cooper, played by Welsh actress Eve Myles, was recruited into Torchwood Three in the series premiere "Everything Changes". A policewoman, partnered with PC Andy Davidson, Gwen by the second series becomes elevated to the team's second-in-command, after serving as leader during Jack's absence. Despite their relationship being tested by Gwen's dishonesty, feelings for Jack Harkness and a tempestuous affair with Owen Harper, Gwen and her live-in boyfriend Rhys eventually get married. After the destruction of the Cardiff branch of the Torchwood Institute, Gwen becomes a mother to daughter Anwen. During Miracle Day she is conflicted between her Torchwood duties and her family responsibilities.